I don't understand this? Just Copella nattereri is listed for Guyana.but i can’t figure out if it’s Marinho or Zarske.
Copella spp. naming is a real mess, because the holotype specimens are too degraded to definitively identify what the fish is.I have some papers on Copella classification if they of any help? One is by Marinho, the other by Zarske?? But I am not sure exactly what you are trying to find out?
According to the pdf you posted there are also C. arnoldi and C. eigenmanni found in Gyuana. (p. 44, 8a and 9a)I don't understand this? Just Copella nattereri is listed for Guyana.